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The disease occurs in Hokuriku and Kanto districts, the central Japan. Sori are produced at ovaries in flowers, and infected flowers show hypertrophy covered with peridium. At first, they are green on the surface, then gradually swollen and turn brown. Approximately 2 mm in size, later the surface disrupts and exposes smut spores (chlamydospores) and then they scatter by wind. Asexual stage is Pseudozyma tsukubaensis (=Candida tsukubaensis) known as an epiphyte on gramineous plants and a producer of erythritol, oligosaccharide.
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